Evolution, Ecology & Organismal Biology

Faculty Listing

EEOB Active Faculty Listing

Participating faculty by last name:    A-C     D-G     H-L     M-R     S-Z

Allen, Edith
Professor of Botany and Plant Sciences
Plant ecology, restoration ecology: Effects of nitrogen deposition on native plant communities, restoration of native vegetation, importance of mycorrhizal fungi in native plant communities.

Allen, Michael (Chair)
Distinguished Professor of Biology,
Professor of Plant Pathology,
Director of the Center for Conservation Biology
Regulation of community and ecosystem processes by soil organisms with special emphasis on mycorrhizal fungi. Global change dynamics and structure of undisturbed areas; conservation and restoration of native ecosystems.

Kurt Anderson
Anderson, Kurt
Assistant Professor of Biology
Quantitative population, community, and applied ecology with an emphasis on modeling spatial dynamics. Responses of organisms to spatial variation in streams and rivers, modeling spatially explicit consumer-resource interactions in terrestrial and aquatic systems, and data-driven modeling for conservation.

Emma Aronson
Aronson, Emma

Assistant Professor of Plant Pathology

Soil microbial ecology and biogeochemistry.


Brelsford, Alan

Assistant Professor of Biology


Hybridization, speciation, and the evolution of sex chromosomes and other supergenes. His research seeks to identify the regions of the genome responsible for evolutionarily important traits, and use variation at regions to study the history of these traits. Current study systems include the population genomics of social behavior in ants, color pattern in birds, sex determination in frogs, and reproductive barriers between related species in all three of these groups. Research in his lab integrates field, lab, and computational methods.


Cardullo, Richard
Professor of Biology and Howard H. Hays Jr. Chair
Biophysics and physiology of fertilization, focusing on molecular interactions between sperm and egg: characterization of egg-associated proteins with complementary receptors on the sperm surface, dynamics of the sperm plasma membrane during fertilization, initiation and characterization of signal transduction pathways leading to the exocytosis of the acrosomal vesicle from sperm.

Chappell, Mark
Professor of Biology
Evolutionary and ecological physiology using a variety of organisms ranging from insects to birds and mammals. Major research topics include adaptation to temperature and high altitude, limits to energy metabolism, and the energy costs of activity in ecologically-relevant contexts.

Clark, Christopher

Assistant Professor of Biology

Courtship displays, the mechanics of how feathers and wings produce sound, and bird flight biomechanics. The relationship between courtship display performance, flight performance, and female preferences, particularly in hummingbirds.
 Matt Daugherty
Daugherty, Matthew
Associate Cooperative Extension Specialist of Entomology
Population biology, pest and disease management, quantitative ecology

Diez, Jeff

Assistant Professor of Plant Ecology

His research is focused on understanding what controls the distributions of species and the composition of communities. These can seem like straight-forward questions but are ultimately really difficult due to the complexity of interacting processes and limitations of available theory and data. Questions about the processes underlying species distributions and community composition are foundational to ecology but have also become more challenging and more urgent in this era of rapid global changes. Climate change and unprecedented rates of species’ movement around the globe (biological invasions) are creating many “novel ecosystems”.  He uses these changes as natural experiments to understand ecosystems, while also building capacity to predict changes in the coming decades.  He also uses field experiments, greenhouse experiments, and statistical modeling that can test hypotheses and quantify processes at different spatial and temporal scales.


Droser, Mary
Professor of Geology
Evolutionary paleoecology, ichnology, the Pre-Cambrian-Cambrian Ordovician radiations, Phanerozoic trends in ecospace utilization, Cambrian and Ordovician of the Great Basin.

 Norman Ellstrand
Ellstrand, Norm
Distinguished Professor of Genetics and Geneticist
The significance of gene flow as an evolutionary force. Applied plant population genetics: (a) gene flow and hybridization as factors in the evolution of increased invasiveness, (b) consequences of unintentional gene flow from domesticated plants to their relatives, and (c) positive and negative impacts of genetically engineered crops, especially with regard to unintentional transgene flow.

Fogel, Marilyn

Professor of Ecology, Wilbur W. Mayhew Endowed Professor of Geoecology, and Director of the EDGE Institute


Stable isotopes of the biologically relevant elements are powerful tools for tracing ecological and geochemical processes on Earth and in planetary materials and environments. Her research for nearly 40 years has concentrated on understanding the flow of elements through modern biogeochemical cycles using stable isotope compositions of organic and inorganic matter as tracers. As she researched elemental cycling in modern ecosystems, she applied this knowledge to understanding how biogeochemical cycles functioned over Earth's history. Her research extends to the fields of paleontology and to astrobiology - searching for evidence of life in the Universe beyond our planet. Inherent in most of the work that she does is the concept of the Earth as a constantly changing entity.


Garland, Theodore, Jr.
Professor of Biology
Evolutionary biology and physiology, with emphasis on the evolution of complex phenotypes; experimental evolution of running behavior and performance in mice; development and application of phylogenetic comparative methods to a variety of evolutionary questions; developoment of free software; lizard and snake locomotor physiology and behavioral ecology.

Gatesy, John
Associate Professor of Biology
Biodiversity and the evolutionary processes that produce it; phylogenetic reconstruction, the inferences that can be made using modern systematic techniques, and development of new methods for the analysis of comparative data.

Hammond, Kim
Professor of Biology, Director of the Natural Reserve System for UCR
Animal physiological ecology and evolutionary physiology, especially the manner in which individuals or species use variation in anatomical and physiological capacities to meet diverse environmental demands.

Hare, Dan
Professor of Entomology 

Evolution and ecology of plant-herbivore and plant-herbivore-natural enemy interactions.

Hayashi, Cheryl
Professor of Biology
The evolution of spider silks across many levels of biological integration, from the molecular genetics of silk genes to protein sequences of different types of silk to biomechanical and functional properties of the final product.

Heraty, John
Professor of Entomology
Morphological and molecular systematics of Chalcidoidea cladistic methodology; biological control evolutionary biology.

Higham, Timothy (Graduate Admissions Advisor)

Assistant Professor of Biology,
Graduate Advisor for Recruitment and Admissions

How animals function mechanically and physiologically in their environments, with emphasis on the biomechanics, muscle physiology and functional morphology of locomotion and feeding in vertebrates. Since physiological mechanisms have been modified over major evolutionary transitions in vertebrate ecology, mechanical analyses are coupled with evolutionary and ecological perspectives.
 Nigel C. Hughes
Hughes, Nigel
Professor of Geology
Field and specimen based approaches to questions of evolutionary mechanism in the early Phanerozoic. Trilobite paleobiology. Lower Paleozoic paleogeography and tectonics (particularly the early Paleozoic history of India and the peri-Gondwanan region), shape restoration of deformed fossils, trace fossil paleobiology, and clastic sedimentology/stratigraphy.

Jenerette, Darrel
Associate Professor of Landscape Ecology
Ecological scaling coupled biogeochemical cycles, terrestrial-aquatic linkages, ecosystem responses to altered precipitation regimes, societal-biophysical interactions.

Litt, Amy

Assistant Professor & Assistant Biologist of Plant Evolution & Development

Our research is aimed at understanding the genetic basis of plant diversity – how genes have changed over the course of evolutionary time, and how those changes have led to the current diversity of plant form and function. We take a comparative interdisciplinary approach, integrating phylogenetics, morphology, and molecular biology methods to characterize changes in gene function across species and how they have influenced plant development.  Current areas of interest include (1) identifying genetic changes that allow some plants to produce edible fleshy fruits, whereas close relatives produce seeds in a dry woody pod; (2) understanding the novel regulatory interactions that occur when two species hybridize, resulting in the combination of two distinct genomes in one nucleus, and often producing unexpected phenotypes; (3) the genetic basis of individual variation in plant development and how selection on this individual variation leads to species diversity; (4) the role of epigenetic factors in domestication, ecological adaptation, and evolution. 


Maduro, Morris
Professor of Biology
Developmental mechanisms of cell fate specification in the nematode C. elegans., particularly the regulatory gene network that specifies the embryonic mesoderm and endoderm precursor cells. The regulators MED-1 and MED-2 function together with the evolutionarily conserved Wnt pathway to specify these cells. Current work involves understanding the molecular basis for Wnt/MED-1,2 activity and elaborating the mesoderm gene network.

Maslov, Dmitri
Professor of Biology
Mitochondrial gene expression in kinetoplastid protozoa, including molecular biology; evolution and parasitology; evolution of kinetoplast DNA and RNA editing; and biodiversity of trypanosomatids, using molecular phylogenetic tools.

Nunney, Len
Professor of Biology
Population and evolutionary genetics, with an emphasis on the application of basic theory to practical problems. Projects include: the population genetics of small conserved populations, the population genetics of cancer, detecting adaptation using genomic data, molecular evolution of Xylella fastidiosa, the role of genetic trade-offs in life history evolution.

Paine, Tim
Professor of Entomology
Biology and ecology of introduced insects in urban environments; interactions of host suitability, host species susceptibility, and natural enemies on insect population biology; pheromone communication systems of bark beetles; interactions between mycorrhizal fungus colonization of plants and the herbivore populations, and the influence of ozone and nitrogen deposition on arthropod communities associated with black oak, ponderosa pine, and bracken fern.

Rafferty, Nicole

Assistant Professor of Biology

Herresearch spans the fields of community ecology, population biology, and global change. She uses experimental manipulations, long-term and historical data, and observations of natural variation to investigate how species interactions are affected by climate change-induced phenological and distributional shifts. These shifts are likely to result in novel communities in the temporal and spatial dimensions, modified interaction strengths, and altered selection on life history events. She is studying these topics in both natural and agricultural plant and pollinator communities.


Rankin, Erin

Assistant Professor of Entomology


Research focuses on investigating species interactions and their effects on trophic dynamics and ecosystem services within the contexts of invasion biology, community ecology and evolutionary ecology. Integrating quantitative, stable isotope and molecular approaches, her work evaluates these interactions by quantifying invader impacts in the field, identifying the mechanisms underlying these interactions, evaluating their contributions to ecosystem services and determining traits that facilitate invasion. Specific areas of current research involve life history evolution of invasive species, invasive generalist predators and trophic impacts of multi-channel omnivory. 


 Khaleel A. Razak
Razak, Khaleel A.
Associate Professor of Psychology
Development of auditory and visual systems, vocalization processing, sound localization and echolocation behaviors, visual motion processing.

Redak, Rick
Professor of Entomology
Plant-insect interactions, conservation biology, community ecology, integrated pest management.

Regan, Helen

Professor of Biology



Quantitative conservation ecology and probabilistic risk assessment, mathematical treatments of uncertainty and decision-making techniques to address conservation management issues.

Reznick, David
Distinguished Professor of Biology
Process of evolution by natural selection explored from an experimental perspective, testing evolutionary theory in natural populations. Guppies from the Caribbean Island of Trinidad are the primary study system, with particular emphasis on the role of predation in the evolution of life history traits, the rate of evolution under natural selection, and the evolution of aging.

Roff, Derek
Professor of Biology
Theoretical and empirical studies of population and quantitative genetics, life-history, and the importance of trade-offs in shaping life history evolution. Current research focuses on insects (especially the importance of trade-offs in determining the evolution of wing dimorphism in various species of crickets) as model systems.

Sachs, Joel

Professor of Biology

Evolution and ecology of symbiotic microbes, evolution of beneficial bacteria & origins of harmful strains, the evolution and breakdown of mutualistic interactions.

Saltzman, Wendy
Professor of Biology
Behavioral endocrinology, especially the bidirectional interactions between hormones and social behavior in mammals. Research emphases include regulation of fertility and endocrine function by the social environment, and interactions between stress and reproductive behaviors.

Santiago, Louis
Associate Professor of Botany and Plant Sciences
Employ a variety of plant physiological techniques, stable isotopes, modeling, phylogenetic analyses, and statistical approaches to understand the ecological implications of the connection between plants and their environment.

Spasojevic, Marko

Assistant Professor of Biology

Plant community ecology, working at the interface of ecology, biogeography, and conservation. Research goals are to understand the mechanisms that influence patterns of biodiversity, and to use that understanding to address environmental issues.

Springer, Mark (Graduate Advisor)

Professor of Biology,
Graduate Advisor for Continuing Students

Molecular evolution and molecular systematics, with an emphasis on the use of molecules to unravel mammalian evolutionary history. Examples include: phylogenetic relationships among the orders of mammals, mammalian molecular clocks, the biogeographic history of mammals in relation to plate tectonic events, reconstructing character transformations for key innovations in mammalian history, the evolution of bats, including the origin of echolocation.

Stajich, Jason

Associate Professor, Department of Plant Pathology and Microbiology and Institute for Integrative Genome Biology

Fungal cell wall evolution, early diverging Chytrid and Zygomycete fungi, post-transcriptional gene regulation, evolution of multicellularity in Fungi, methods in comparative and evolutionary genomics, human pathogenic fungi.
Walton, Bill
Professor of Entomology
IPM of vector and pest arthropods particularly mosquitoes, biogeography of freshwater flora and fauna, trophic interactions of freshwater food webs.

Bradley White
White, Bradley 

Adjunct Assistant Professor of Entomology

Genomics of malaria mosquitoes with an emphasis on understanding the genetic basis of medically-relevant phenotypic diversity.  Population, ecological, and quantitative genomics approaches are combined with field work and experimental genetics to connect genotype to phenotype and investigate the origin and maintenance of functional nucleotide polymorphisms.


Woodard, Hollis S.

Assistant Professor of Entomology


Ecology, evolution, and conservation of native bees, with a focus on bee nutritional ecology and feeding biology.


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Evolution, Ecology and Organismal Biology (EEOB) Graduate Program
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